Structure is not mechanism: high-gain gated-FFN rows across text and genomic foundation models

The study investigates whether unusually high‑gain parameters in transformer models—specifically gated feed‑forward network (gated‑FFN) rows—play a critical functional role across both text and genomic foundation models. By computing exact bilinear weight operators and testing structural extremeness, the authors find that high‑gain rows are enriched for functional importance but do not reliably predict causal effect size or severity. The analysis reveals model‑specific causal organizations, including super‑additive interactions in DNABERT‑2 and position‑localized dependencies in GENERator, indicating that structural prominence signals enrichment rather than calibrated criticality.

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